Showing posts with label Wildlife Photography. Show all posts
Showing posts with label Wildlife Photography. Show all posts

Common Greenshank (Tringa nebularia): Taxonomy, Identification, Ecology and Occurrence at Mahanadi River at Cuttack, Odisha,

 Common Greenshank (Tringa nebularia): Taxonomy, Identification, Ecology and Occurrence at Chilika Lake, Odisha

Abstract

The Common Greenshank (Tringa nebularia) is a widespread Palearctic migratory shorebird belonging to the family Scolopacidae. It breeds in the boreal and subarctic wetlands of northern Europe and Asia and winters across Africa, southern Asia, Southeast Asia, and Australasia. During the non-breeding season, the species frequents a wide variety of freshwater and coastal wetlands. The present note documents an individual observed along the Mahanadi River at Cuttack, Odisha, during April. Morphological characteristics visible in the photographs—including the long, slightly upcurved bill, greenish legs, robust body structure, and non-breeding plumage—confirm its identity as the Common Greenshank. This account reviews the species' taxonomy, recent systematic position, identification, ecology, migration, and significance within the freshwater riverine ecosystem of eastern India.


Introduction

The Mahanadi River is one of eastern India's major river systems and supports an extensive mosaic of sandbars, shallow channels, exposed mudflats, and seasonally inundated islands. These habitats provide important feeding and resting grounds for numerous migratory waterbirds arriving from northern Eurasia during the winter months.

Although Odisha is internationally recognized for the avian diversity of Chilika Lake, the riverine wetlands of the Mahanadi basin also constitute significant habitats for migratory shorebirds. Species of the family Scolopacidae, particularly members of the genus Tringa, regularly utilize these inland freshwater environments during migration and winter. Among them, the Common Greenshank (Tringa nebularia) is one of the largest and most conspicuous waders encountered on river sandbars and shallow margins.

The individual documented here was photographed standing in shallow flowing water on the Mahanadi River near Cuttack during April, representing a late spring migrant prior to its northward departure.

Tringa nebularia from Odisha observed April 2025


Taxonomic Classification

Kingdom: Animalia

Phylum: Chordata

Class: Aves

Clade: Neoaves

Order: Charadriiformes

Suborder: Scolopaci

Family: Scolopacidae

Subfamily: Tringinae

Genus: Tringa

Species: Tringa nebularia (Gunnerus, 1767)


Recent Systematic Position

The systematic arrangement of shorebirds has undergone considerable revision over the past two decades due to advances in molecular phylogenetics.

Earlier classifications were primarily based on external morphology, skeletal anatomy, plumage, and feeding behavior. Under these traditional systems, the Common Greenshank was placed within the genus Tringa along with shanks, tattlers, and yellowlegs.

Recent molecular studies employing mitochondrial DNA sequences, nuclear genes, and genome-scale analyses have confirmed that:

  • The family Scolopacidae is a monophyletic lineage within the order Charadriiformes.

  • The genus Tringa forms a well-supported evolutionary clade.

  • The Common Greenshank is evolutionarily closest to the Greater Yellowlegs (Tringa melanoleuca) and Lesser Yellowlegs (Tringa flavipes), rather than to superficially similar species such as the Marsh Sandpiper.

  • The Marsh Sandpiper (Tringa stagnatilis) diverged earlier within the Tringa lineage despite possessing convergent external morphology.

Recent genomic studies further indicate that the traditional "shanks" comprise a distinct evolutionary radiation within the subfamily Tringinae, characterized by long-legged wading adaptations and diverse foraging strategies.



The currently accepted classification adopted by the International Ornithologists' Union (IOC), the Clements Checklist, and the BirdLife International Checklist continues to recognize Tringa nebularia as a valid species without subspecies.


Diagnostic Morphology

The photographed bird exhibits all major diagnostic characters of Tringa nebularia.

General appearance

The Common Greenshank is a relatively large wader measuring:

  • Length: 30–35 cm

  • Wingspan: 55–70 cm

  • Weight: 130–290 g

The species possesses an elongated body supported by long greenish legs adapted for wading in deeper water than most sandpipers.

Bill

Perhaps the most diagnostic feature is the bill.

The bill is:

  • Long

  • Moderately stout

  • Dark grey to black

  • Slightly upturned near the tip

This gentle upward curvature is clearly visible in the second photograph and immediately distinguishes the bird from numerous similar species.

Plumage

During the non-breeding season the plumage consists of:

  • Pale grey upperparts

  • Finely vermiculated scapular feathers

  • White belly

  • Light streaking over the breast

  • White supercilium

  • Distinct black-and-white barring on the tail

The observed individual perfectly matches this plumage stage.


Identification from the Photographs


Several field marks support the identification.

1. Slightly upturned bill

The bill is longer and heavier than that of Marsh Sandpiper and curves gently upward.

2. Long green legs

Unlike yellowlegs, whose legs are bright yellow, the legs here are dull greenish-grey.

3. Robust body

The bird appears compact and muscular rather than delicate.

4. Tail barring

The black-and-white tail barring visible in the first image is typical of Common Greenshank.

5. Habitat preference

Standing in relatively deep shallow water is characteristic of Greenshanks, which often forage in deeper water than smaller sandpipers.


Comparison with Similar Species

Marsh Sandpiper (Tringa stagnatilis)

Although frequently occurring at Chilika, Marsh Sandpiper differs by possessing:

  • extremely slender body

  • needle-like straight bill

  • proportionately much longer legs

  • more graceful posture

  • lighter head pattern

The photographed individual lacks these characters.

Common Redshank (Tringa totanus)

Redshanks possess:

  • bright orange-red legs

  • orange bill base

  • heavier breast streaking

Neither feature is present.

Wood Sandpiper (Tringa glareola)

Wood Sandpipers are:

  • considerably smaller

  • heavily spotted above

  • possess a conspicuous pale eyebrow

Again, these features are absent.


Feeding Ecology

The Common Greenshank is an opportunistic carnivorous feeder.

Diet includes:

  • polychaete worms

  • aquatic insect larvae

  • molluscs

  • shrimps

  • crabs

  • small fishes

  • tadpoles

Unlike many sandpipers that employ rapid pecking, Greenshanks frequently sweep their bills laterally through shallow water while walking.



Breeding Biology

Breeding occurs in northern Eurasia from May through July.

The nest is a shallow depression on open tundra near wetlands.

Typical clutch:

  • 4 eggs

  • incubation approximately 24 days

  • chicks are precocial

  • fledging occurs after about one month

The species exhibits strong territorial behaviour during breeding but becomes highly gregarious on migration.


Migration

The Common Greenshank undertakes long-distance migration along several major flyways.

Indian birds primarily belong to populations breeding in:

  • Scandinavia

  • Finland

  • Russia

  • Siberia

These populations migrate south via the Central Asian Flyway, arriving in India between August and October.

Northward migration begins during March and April, making the present April observation entirely consistent with the species' annual migratory schedule.

Occurrence in Odisha

In Odisha, the Common Greenshank is a regular winter visitor and passage migrant. While frequently reported from Chilika Lake, the species is equally recorded from inland wetlands, including the Mahanadi river system, Hirakud Reservoir, Ansupa Lake, Bhitarkanika estuaries, and numerous floodplain wetlands. April records represent birds on northward migration to their breeding grounds in northern Eurasia.


Conservation Status

IUCN Red List: Least Concern (LC)

Although globally stable, local populations face threats from:

  • wetland degradation

  • pollution

  • habitat reclamation

  • disturbance by tourism

  • climate change affecting migration timing

Protection of internationally important wetlands such as Chilika remains essential for maintaining migratory shorebird populations.


Conclusion

The bird photographed at Mahanadi River at Cuttack, Odisha, in April is confidently identified as the Common Greenshank (Tringa nebularia) based on its slightly upturned robust bill, long greenish legs, overall structure, non-breeding plumage, and characteristic tail barring. Modern molecular phylogenetic studies support its placement within the monophyletic genus Tringa in the subfamily Tringinae, where it is most closely related to the yellowlegs (T. melanoleuca and T. flavipes). Its presence at Chilika underscores the ecological importance of this Ramsar wetland as a critical stopover and wintering site along the Central Asian Flyway.


Selected References

  • Gill, F., Donsker, D. & Rasmussen, P. (eds.). IOC World Bird List (current version).

  • del Hoyo, J., Elliott, A., & Christie, D. A. (eds.). Handbook of the Birds of the World. Lynx Edicions.

  • Gibson, R. & Baker, A. J. (2012). Multiple gene sequences resolve phylogenetic relationships within the shorebirds (Scolopacidae). Molecular Phylogenetics and Evolution.

  • Piersma, T., Wiersma, P. & Kirwan, G. M. Birds of the World: Common Greenshank. Cornell Lab of Ornithology.

  • Praveen, J., Jayapal, R. & Pittie, A. Checklist of the Birds of India (latest edition).

Dr. U Elaya Perumal

The Silent Sentinel of a River: A Field Encounter with the Striated Heron (Butorides striata) in Central India

 The Silent Sentinel of a River: A Field Encounter with the Striated Heron (Butorides striata) in Central India

Abstract

Not every remarkable wildlife encounter occurs inside a national park or a well-known bird sanctuary. Sometimes, a small freshwater rivulet hidden among rocks and riparian vegetation reveals an equally fascinating story of survival. During a field visit on 20 April 2025, a solitary Striated Heron (Butorides striata) was observed hunting along a rivulet immediately upstream of its confluence with the Satna River in Madhya Pradesh, India (24.507557° N, 80.875699° E). The observation offered an opportunity to study one of India's most accomplished freshwater predators in its natural environment.


Unlike conspicuous egrets that often dominate open wetlands, the Striated Heron relies on camouflage, patience, and precision rather than speed. This article explores the natural history of the species through a field observation, highlighting how even small river systems contribute to freshwater biodiversity and why protecting such habitats is essential for sustaining resident bird populations.


Where Two Waters Meet

Every river has a story, and many begin long before the main channel becomes visible on a map. Small rivulets, seasonal streams, and rocky channels quietly gather rainwater and groundwater, carrying life toward larger rivers. These seemingly modest watercourses are often overlooked, yet they support a surprising diversity of aquatic organisms and wildlife.

On 20 April 2025, while walking along one such rivulet just before it merged with the Satna River in central India, the landscape appeared almost motionless. Clear water flowed gently between weathered rocks, dragonflies skimmed the surface, and the calls of distant birds echoed through the riparian vegetation. At first glance, nothing suggested the presence of a predator.

Then a slight movement along the opposite bank caught the eye.

Standing perfectly still among stones and shallow water was a bird so well camouflaged that it almost disappeared into the landscape. Only its bright yellow eye betrayed its presence. The bird remained motionless for several minutes before taking a slow, deliberate step toward the water's edge. A few seconds later, it struck with astonishing speed.

The hunter was a Striated Heron (Butorides striata).


An Artist of Disappearance

Unlike larger members of the heron family that often feed in open marshes or paddy fields, the Striated Heron specializes in discretion. Every aspect of its appearance appears designed to avoid detection.


The slate-grey crown resembles wet river stones. Olive-brown wings blend with weathered rock and dried vegetation. Fine streaks across the throat mimic the play of light and shadow along the riverbank. Even when standing only a few metres away, the bird can remain almost invisible.

This extraordinary camouflage is not merely defensive—it is an essential hunting adaptation.

Many freshwater fishes, tadpoles, and aquatic insects are highly sensitive to movement. Any sudden motion may send potential prey darting into deeper water. Remaining undetected gives the heron a decisive advantage, allowing it to approach prey without triggering an escape response.

The photographs accompanying this observation beautifully illustrate this adaptation. Against the rocky substrate of the rivulet, the bird's plumage merges almost seamlessly with its surroundings, demonstrating why casual observers often overlook this species despite its widespread distribution across India.


A Hunter That Measures Time Differently

Watching the Striated Heron challenges our usual perception of hunting. Modern predators are often associated with speed—a falcon diving through the sky or a kingfisher plunging into water. The Striated Heron follows a different philosophy.

It hunts through patience.

For several minutes, the bird scarcely moved. Its body remained low, neck partially folded, eyes fixed upon the shallow current. Rather than walking continuously, it advanced one cautious step at a time, pausing repeatedly as though studying every ripple in the water.

When prey finally entered striking distance, the attack was almost impossible to follow with the naked eye. The neck unfolded in a fraction of a second, transforming the apparently motionless bird into an extraordinarily efficient predator.

This combination of stillness and explosive speed has enabled the Striated Heron to thrive in freshwater ecosystems across much of the world.


Identifying the Observer's Bird

The individual observed displayed the diagnostic characteristics of an adult Striated Heron (Butorides striata), including:

  • Slate-grey crown and mantle.
  • Rich chestnut-brown sides of the neck.
  • White throat marked with fine dark streaks.
  • Bright yellow iris.
  • Greenish-yellow lores.
  • Long, sharply pointed black bill.
  • Yellowish-green legs.
  • Pale spotting on the wing coverts.
  • Compact body with a characteristic crouched hunting posture.

These features distinguish it from superficially similar species such as the Indian Pond Heron (Ardeola grayii), which has heavier streaking on the neck and a different body profile.




More Than a Bird

This encounter was also a reminder that biodiversity is not confined to protected reserves or internationally recognized wetlands. Small tributaries, seasonal rivulets, and river confluences serve as vital habitats for countless species, many of which remain largely unnoticed.

Healthy freshwater ecosystems are built upon these smaller waterways. They support aquatic insects, fishes, amphibians, reptiles, mammals, and birds, forming ecological networks that ultimately sustain larger river systems.

The solitary Striated Heron standing silently beside this rivulet represented not only a remarkable bird but also the ecological integrity of the stream itself.

The Master of Stillness

The first impression of a Striated Heron is often deceptive. Standing motionless along the water's edge, it appears almost detached from its surroundings, as though it were simply another weathered stone on the riverbank. Yet beneath this calm exterior lies one of the most efficient freshwater predators found in Asian rivers and wetlands.

Unlike many birds that rely on speed or endurance, the Striated Heron has evolved an entirely different hunting strategy—one based on patience, precision, and an exceptional understanding of its environment.


The Art of Ambush

Every movement made by the Striated Heron appears deliberate. During the observation at the rivulet flowing into the Satna River, the bird remained almost perfectly still for several minutes. Only subtle head movements indicated that it was continuously monitoring the shallow water before it.

Rather than actively chasing prey, the heron allows prey to come within striking distance. When a small fish, tadpole, or aquatic insect moves close enough, the folded neck suddenly extends with remarkable speed, driving the sharp bill into the water with extraordinary accuracy.

This ambush strategy offers several advantages. It conserves energy, minimizes disturbance to the water surface, and increases the likelihood of catching alert prey that would quickly escape from a more active hunter. In clear, shallow streams where prey can detect movement easily, patience often proves more effective than speed.


Built for Life Along the Water

The Striated Heron possesses several anatomical adaptations that make it exceptionally suited to hunting in freshwater habitats.

Its long, dagger-shaped bill functions as both a spear and forceps, allowing the bird to seize slippery prey with precision. The neck contains highly flexible vertebrae and powerful muscles that enable the rapid extension necessary for striking.

Equally important are its eyes. Positioned to provide excellent binocular vision, they allow the heron to judge distances accurately despite the refraction of light through water. This depth perception is essential when targeting fast-moving prey beneath the surface.

Its relatively short legs, compared with larger herons, are ideal for navigating rocky river margins, submerged stones, and shallow pools. The compact body and low centre of gravity also improve balance on uneven substrates, enabling the bird to hunt effectively where larger wading birds may struggle.


A Diverse Freshwater Diet

The Striated Heron is an opportunistic carnivore whose diet reflects the diversity of organisms found in healthy freshwater ecosystems.

Its principal prey includes:

  • small fishes,
  • tadpoles,
  • frogs,
  • freshwater shrimps,
  • crabs,
  • aquatic beetles,
  • dragonfly larvae,
  • water bugs,
  • molluscs,
  • small reptiles, including young lizards.

The exact composition of the diet varies with habitat and season. In shallow streams such as the one observed in central India, fishes and aquatic insects form the bulk of its prey, while crabs and shrimps become increasingly important in estuarine and mangrove environments.

Because the species occupies a relatively high position in the aquatic food web, it also serves as an indicator of healthy freshwater ecosystems. Rivers capable of sustaining Striated Herons generally support diverse fish populations and productive invertebrate communities.


One of Nature's Patient Hunters

Among freshwater birds, few display the remarkable patience of the Striated Heron. Individuals have been observed remaining motionless for extended periods before making a single decisive strike.

This behaviour requires exceptional energy efficiency. Instead of expending energy through constant movement, the bird invests time in careful observation. Every ripple, shadow, and flash beneath the water's surface is assessed before action is taken.

The field photographs captured during this observation illustrate this perfectly. In both images, the bird adopts its characteristic low, forward-leaning posture with the neck partially folded, ready to strike without warning. Such behaviour exemplifies why the species is regarded as one of the finest ambush predators among freshwater birds.


A Bird That Uses Tools

Perhaps one of the most remarkable aspects of the Striated Heron's natural history is its documented ability to use tools—an uncommon behaviour among birds.

Researchers have observed individuals placing feathers, leaves, insects, or small twigs on the water surface to attract curious fish. When a fish approaches the floating object, the heron rapidly captures it.

This behaviour has been documented in several parts of the world and demonstrates a level of behavioural flexibility rarely associated with wading birds. Although not every individual uses tools, the observations suggest that the species possesses considerable learning ability and can adapt its hunting techniques to local conditions (Lefebvre et al., 2002).


Life Along Small Rivers

Large wetlands often receive considerable conservation attention, but smaller streams and rivulets are equally important for species such as the Striated Heron.

The unnamed rivulet observed during this field visit, flowing into the Satna River, illustrates this point well. Its shallow pools, gently flowing water, rocky margins, and patches of riparian vegetation create a mosaic of microhabitats supporting fish, amphibians, aquatic insects, and crustaceans. These organisms, in turn, sustain predators like the Striated Heron.

Such small watercourses also function as ecological corridors, linking larger rivers and wetlands while facilitating the movement of aquatic organisms. Protecting these tributaries is therefore essential for maintaining the overall health of river ecosystems.


An Indicator of Healthy Freshwater Ecosystems

Although the Striated Heron is classified globally as a species of Least Concern, its continued presence depends upon clean, functioning freshwater habitats.

Healthy rivers provide:

  • abundant prey,
  • stable water levels,
  • natural riverbank vegetation,
  • undisturbed nesting sites,
  • minimal chemical pollution.

Conversely, declining water quality, excessive sand mining, channel modification, and the removal of riparian vegetation reduce habitat suitability for the species. Because of its close association with freshwater ecosystems, the Striated Heron is often regarded as a useful indicator of river health.

Observing this bird along a relatively undisturbed tributary of the Satna River serves as a reminder that even modest streams can harbour rich biodiversity when their ecological integrity is maintained.

Rivers Worth Protecting

The quiet observation of a Striated Heron along a small rivulet before it joins the Satna River is more than an interesting bird sighting. It is a reminder that healthy freshwater ecosystems are built from countless small streams and tributaries that collectively sustain biodiversity across an entire river basin. Although these habitats rarely receive the attention given to major rivers or protected wetlands, they are indispensable for wildlife and human communities alike.


A Resident of Rivers Across the World

The Striated Heron (Butorides striata) is among the most widely distributed members of the heron family. Its range extends across Africa, Madagascar, southern Europe, the Middle East, South Asia, Southeast Asia, Australia, and numerous Pacific islands (BirdLife International, 2024).

In India, it is a resident species, occurring throughout the year in a wide variety of freshwater and coastal habitats. It is frequently encountered along:

  • rivers and tributaries,
  • lakes and reservoirs,
  • irrigation canals,
  • village ponds,
  • marshes,
  • mangroves,
  • estuaries,
  • paddy fields.

Unlike many migratory waterbirds, the Striated Heron does not undertake long seasonal migrations. Instead, individuals usually remain within local river systems, moving only short distances in response to changing water levels or food availability.

Because of this close association with freshwater habitats, local populations often reflect the ecological condition of the rivers they inhabit.




Breeding Along Quiet Waterways

Breeding generally coincides with the onset of the monsoon or periods of increased food availability, although timing varies across different parts of India.

The Striated Heron usually nests in secluded places where disturbance is minimal. Nests are commonly built in:

  • riverside trees,
  • shrubs overhanging water,
  • dense riparian vegetation,
  • mangroves,
  • bamboo clumps.

The nest itself is a simple platform of twigs lined with smaller plant material.

Typical breeding characteristics include:

  • clutch size of 3–5 pale blue-green eggs,
  • incubation lasting about 20–23 days,
  • both parents sharing incubation and chick rearing,
  • nestlings remaining in the nest for approximately three weeks before fledging.

The dependence on riverside vegetation highlights the importance of conserving natural riparian habitats. Removal of trees and shrubs along riverbanks not only accelerates erosion but also eliminates suitable nesting sites for many bird species.


Challenges Facing Freshwater Birds

Although the Striated Heron is currently listed as Least Concern by the International Union for Conservation of Nature (IUCN), its habitat is increasingly affected by human activities (BirdLife International, 2024).

Major threats include:

  • pollution from untreated sewage,
  • agricultural runoff,
  • indiscriminate sand mining,
  • river channel modification,
  • removal of riparian vegetation,
  • construction activities,
  • excessive extraction of river water,
  • plastic pollution,
  • disturbance from human recreation.

Smaller tributaries are particularly vulnerable because they are often excluded from formal conservation programmes despite supporting diverse aquatic communities.

The cumulative impact of habitat degradation may not immediately eliminate resident birds, but it gradually reduces prey availability, nesting opportunities, and overall ecosystem resilience.


Why Small Tributaries Matter

Large rivers depend upon healthy tributaries. Every stream contributes water, nutrients, sediments, and biodiversity to the main river channel.

The rivulet where this Striated Heron was photographed illustrates this principle perfectly. Despite its modest size, it supports aquatic insects, fishes, amphibians, reptiles, and birds, functioning as a self-contained ecosystem before merging with the Satna River.

Such habitats provide:

  • breeding areas for fishes,
  • refuges for amphibians,
  • feeding grounds for waterbirds,
  • corridors connecting larger aquatic ecosystems,
  • natural filtration of sediments and nutrients.

Protecting these smaller waterways therefore benefits the entire river basin.

The presence of a top freshwater predator such as the Striated Heron suggests that the ecosystem still retains much of its natural ecological complexity.


A Bird That Represents Healthy Rivers

Birds often serve as ambassadors for ecosystem conservation, and the Striated Heron is no exception.

Unlike spectacular migratory flocks that attract immediate attention, this species quietly performs its ecological role every day by regulating populations of fishes, insects, amphibians, and crustaceans. Its survival depends on clean water, abundant prey, and undisturbed riverbanks.

For naturalists, photographers, and conservationists, encountering a Striated Heron is often a sign that a river still possesses many of the characteristics of a functioning freshwater ecosystem.

Each observation therefore contributes valuable information to our understanding of local biodiversity and highlights the importance of continued monitoring of India's inland waterways.


Conclusion

The observation of a Striated Heron (Butorides striata) on 20 April 2025 along a small freshwater rivulet immediately upstream of its confluence with the Satna River in Madhya Pradesh demonstrates how even modest watercourses support remarkable wildlife.

Patient, inconspicuous, and exceptionally well adapted to its environment, the Striated Heron exemplifies the intricate relationships that exist within freshwater ecosystems. Its camouflage, hunting strategy, and dependence on healthy aquatic habitats make it one of the most fascinating resident birds of India's rivers.

The encounter also underscores an important conservation message: safeguarding biodiversity is not limited to protecting large wetlands or famous rivers. Countless unnamed streams, rivulets, and tributaries form the ecological foundation of larger river systems. Their preservation ensures the survival of species like the Striated Heron and countless other organisms that depend upon clean, flowing freshwater.

As pressures from pollution, habitat alteration, and unsustainable resource use continue to grow, protecting these overlooked waterways will become increasingly important. Every healthy tributary strengthens the resilience of the larger river it joins, reminding us that conservation often begins in the smallest places.


References (APA 7th Edition)

Ali, S. (2002). The Book of Indian Birds (13th ed.). Bombay Natural History Society.

BirdLife International. (2024). Butorides striata (Striated Heron). BirdLife Data Zone.

del Hoyo, J., Elliott, A., Sargatal, J., Christie, D. A., & de Juana, E. (Eds.). (2020). Handbook of the Birds of the World Alive. Lynx Edicions.

Gill, F., Donsker, D., & Rasmussen, P. (Eds.). (2024). IOC World Bird List (Version 14.1). International Ornithologists' Union.

Grimmett, R., Inskipp, C., & Inskipp, T. (2011). Birds of the Indian Subcontinent (2nd ed.). Oxford University Press.

Kushlan, J. A., & Hancock, J. A. (2005). The Herons. Oxford University Press.

Rasmussen, P. C., & Anderton, J. C. (2005). Birds of South Asia: The Ripley Guide. Smithsonian Institution & Lynx Edicions.

Wetlands International. (2023). Waterbird Conservation in Asia and the Central Asian Flyway.

by Dr. U Elaya Perumal.

Common Greenshank (Tringa nebularia): Taxonomy, Identification, Ecology and Occurrence at Mahanadi River at Cuttack, Odisha,

  Common Greenshank ( Tringa nebularia ): Taxonomy, Identification, Ecology and Occurrence at Chilika Lake, Odisha Abstract The Common Green...